AFTOL: Resolving the Evolutionary History of the Fungi is a project that involves researchers at ten universities and many collaborators in the mycological community. It will contribute to understanding the lineage of fungi---fungi phylogeny---at multiple levels and will promote training and education in fungal biology. In support of this project, the research group of Professor David J. McLaughlin (Plant Biology) is coordinating analysis of subcellular characters as well as studying molecular phylogenetics of selected species. Part of this project includes the development of a Structural and Biochemical Database. This database will become a major resource for the scientific community, managing and providing morphological and biochemical information on fungi and serving as a phyloinformatics tool.